- In amphibians blood flow is directed in two circuits: one
through the lungs and back to the heart, which is called
pulmonary circulation, and the other throughout
the rest of the body and its organs including the brain (systemic
circulation). In amphibians, the gas exchange also occurs through
the skin during pulmonary circulation and is referred to as
pulmocutaneous circulation. amphibians have a
three-chambered heart that has two atria and one ventricle. The two
atria (superior heart chambers) receive blood from
the two different circuits (the lungs and the systems), and then
there is some mixing of the blood in the heart’s
ventricle (inferior heart chamber), which reduces
the efficiency of oxygenation. The advantage of this arrangement is
that high pressure in the vessels pushes blood to the lungs and
body. The mixing is mitigated by a ridge within the ventricle that
diverts oxygen-rich blood through the systemic circulatory system
and deoxygenated blood to the pulmocutaneous circuit. For this
reason, amphibians are often described as having double
circulation.
- Amphibians have evolved multiple ways of breathing. Young
amphibians, like tadpoles, use gills to breathe, and they do not
leave the water. As the tadpole grows, the gills disappear and
lungs grow. A large surface area to allow as much oxygen to enter
the gills as possible because more of the gas comes into contact
with the membrane. he other means of breathing for amphibians is
diffusion across the skin. To aid this diffusion, amphibian skin
must remain moist. It has vascular tissues to make this gaseous
exchange possible. This moist skin interface can be a detriment on
land but works well underwater.
COMPARISION OF amphibian respiratory and circulatory
system structure to human respiratory and circulatory system
structures
Circulatory System
Amphibians
- Most have three chambers; two atria and one ventricle.
- The ridge in the ventricle takes most of the oxygen-poor blood
from the right atrium to the pulmocutaneous circuit and most of the
oxygen-rich blood from the left atrium to the systemic circuit
- When the amphibian submerges in water, they shut off their
blood flow to their lungs and continues blood flow to the skin for
gas exchange.
Humans
- Have two atria and two completely divided ventricles. The left
side of the heart pumps and receives only oxygen-rich blood while
the right side of the heart pumps and receives only oxygen-poor
blood.
- Humans have typically bigger and take up more energy so their
circulatory systems need to deliver more oxygen to their tissues.
Humans can do this by their separate and independent systemic and
pulmonary circuits.
Respiratory System
Amphibians
Adult amphibians are lung-breathers. The skin acts as an
accessory respiratory organ both in water and on land. The skin is
highly vascular and specially so in the buccopharyngeal
cavity.
Few urodeles retain external gills as the respiratory organs in
adults.
Both external and internal gills are present in anuran larvae.
Ascaphus, living in the mountain stream of U.S.A., has reduced
lungs which help the animal to live in water.
In almost all amphibians cutaneous respiration is a remarkable
supplementary respiratory adaptation.
In caecilians, the tracheal lung may be present but the left one
is always rudimentary. In aquatic urodeles, the lungs act
secondarily as hydrostatic organ. In all these above cases,
respiration is exclusively pharyngeal and/or cutaneous.
Humans
The primary organs of the respiratory system are the lungs,
which carry out this exchange of gases as we breathe
As we breathe, oxygen enters the nose or mouth and passes the
sinuses, which are hollow spaces in the skull that help regulate
the temperature and humidity of the air we breathe.
The alveolar walls are extremely thin (about 0.2 micrometers)
and are composed of a single layer of tissues called epithelial
cells and tiny blood vessels called pulmonary capillaries. Blood in
the capillaries picks up oxygen and drops off carbon dioxide.
The diaphragm, a dome-shaped muscle at the bottom of the lungs,
controls breathing and separates the chest cavity from the
abdominal cavity.